The index is calculated using the formula:
In this formula, represents the total number of individuals of a single species, while represents the total number of individuals of all species combined.
The term represents the probability that two individuals randomly selected from the sample will belong to the same species. By subtracting the sum of these probabilities from 1, we find the probability that two individuals belong to different species.
Step 1: Data Collection: Count the number of individuals () for every species present in the sample area and sum them to find the total population ().
Step 2: Relative Abundance: For each species, divide its population by the total population to find the proportion .
Step 3: Squaring: Square each of the resulting proportions. This step gives more weight to the most common species in the calculation.
Step 4: Summation and Subtraction: Add all the squared values together and subtract this total from 1 to arrive at the final value for .
| Feature | Species Richness | Species Evenness |
|---|---|---|
| Definition | Total count of different species. | Relative abundance of each species. |
| Sensitivity | Ignores population sizes. | Highly sensitive to population distribution. |
| Role in Index | Increases as more species are added. | Increases as populations become more balanced. |
Check the Formula: Always ensure you are using the correct version of the formula provided. Some contexts use while others use ; always follow the specific notation required by your syllabus.
The 'Subtract from 1' Rule: A very common mistake is calculating and stopping there. Remember that this value represents 'dominance'; you must subtract it from 1 to find 'diversity'.
Rounding Errors: Carry at least three or four decimal places through your intermediate steps (the squaring and summing) to avoid significant rounding errors in the final index value.
Sanity Check: If your final answer is greater than 1 or less than 0, a calculation error has occurred. Re-check your division and summation steps immediately.